A long time ago in a bog far, far away...

Posted by Graeme Lyons , Saturday, 7 March 2020 08:55

Well about an hours drive away. Just before I left my seven years at the RSPB, one of the last thigs I did was a load of monitoring at Broadwater Warren RSPB reserve, a recently acquired conifer plantation with some isolated pockets of heath, acid-grassland and mire. I did a baseline NVC survey but also put out a series of pitfall traps and pan traps in the existing habitat. I didn't know how to identify any of this stuff back than, 12.5 years ago. The spiders were passed to Ian Dawson but because the records were in the SxBRC, when I set up my database, the records were synced with my database. 

So I have frustratingly a number of records for spiders that I have technically seen but don't feel like I have and won't count them on my own personal list. Including some goodies, like the only Sussex record of Jacksonella falconeri, our smallest spider and loads of records of Walckenaeria furcillata, what must be my most wanted spider. All in all, there were 82 species recorded and 8 of these have status (a respectable 9.8%). I am sure that more spiders have been recorded since but I sued this as the site species list and headed back there yesterday. Probably my only day out in March.

The posts from pan traps are almost all still in place, despite so much grazing! Even a bit of solidified raspberry net was still in place, what I used to pin the traps down.

I recorded 34 species yesterday in half a day. Most of the action was in the bog. A lifer for me was this diminutive Hahnia pusilla (NS) sieved from Sphagnum. Only the 2nd Sussex record, and the first since 1969.

I was struck by how many of the species recorded yesterday were not picked up in the pitfalls in 2007, albeit they were in the summer so it's not that surprising. All in all 21 new species and six of these had status, bringing the records I have for the site to 103 and 14 with status (13.6%), suggesting the assemblage is improving, which marries up with observation on site.

The other goody was my first Sussex Walckenaeria nodosa, and the first in Sussex since 1969 also! The other new species with status were Cercidia prominens, Sintula corniger and the ubiquitous Meioneta mollis.

I went to have a look at this W5 Alder carr, I was expecting great thing from the tussocks but for some reason they were not very rich but I did get Theridiosoma gemmosum from them.

The huge changes on this site though are the great expanses of open heathland, all this was under conifers 12.5 years ago, a remarkable achievement. Woodlark were singing away nearby.

It's amazing how much things have changed for me in 12.5 years, or an eighth of a century. I was so desperate to move back to Brighton then living in Cambridge at the time and moving back down here and getting the job at the Trust was an amazing experience. Once upon a time I never thought I'd leave the RSPB, then I thought I'd never leave the Trust! I never thought then though that I'd end up as a freelance entomologist and county recorder for various groups. Pan-species listing wasn't even a thing then. I was in my late twenties when I last looked at this site in detail and now I'm in my forties, about to start the next chapter. 

I absolutely can't wait for my first full year of freelance, I can't wait to get out in the field too! My spider year list is now on 160 with nine new species today. I will try and go back in the summer to refind Walckenaeria furcillata.

Leap frog

Posted by Graeme Lyons , Saturday, 29 February 2020 21:34

Once every 1461 days we get a 29th February and a chance to see a 'leap frog' (this isn't a thing, I made it up). By a total coincidence I did exactly that today and got a new amphibian thanks to James Harding-Morris, Seth Gibson and Mark Telfer. I had a long overdue day off and headed to the butterfly house again at Whipsnade Zoo. All that will have to wait as it was rather eclipsed by these beauties at the end of the day in Bedfordshire. This is the Midwife Toad Alytes obstetricans. What a smart little beast and a great scientific name too. Introduced but apparently not a major threat being a poor disperser. James found three very quickly and I indulged in some shots.

Such incredible eyes on them. The rather sickly colour and texture of the skin reminds me of the zombies from the Walking Dead! Such pleasing proportions too and great to see them next to Common Toad. This little one really showed the lines of red warts that are really distinctive.
I didn't get to hear the Call of the Midwife Toad, (which sounds like a bar code scanner), but you can hear them on this video here on this One Show piece from 2010. Thanks all for a great day!

Emarginal gains

Posted by Graeme Lyons , Friday, 31 January 2020 14:34

Yes I did just make a word up as a tenuous attempt at a pun. Live with it. Last November I went spidering to the New Forest with Mark Gurney to a National Trust site called Ibsley Common. We found some nice things that had not been seen in the Forest for some time, such as Hypselistes, Taranuncus and Notioscopus. All nice bog species but not much that I hadn't already seen that year. In fact my only year tick I think was Oedothorax fuscus. But I did sweep a smart looking immature Philodromus from Bog Myrtle in a bog under some pines. I became convinced that it was going to be Philodromus emarginatus despite that the fact that I had never seen it. Maybe it was the large flock of Hawfinches that just made you feel like everything there was going to be rare but I had a nagging hunch that I couldn't shift.

Here is the photo of I took in the field at the end of November.

I consulted a few people but knew what the answer was going to be. I needed to get it to adulthood. Further to this Peter Harvey said "I can see nothing much to see emarginatus is a strong contender". So, I HAD to get it to maturity, as I really felt like this was very different to all the Philodromus I had seen before, including the washed out margaritatus that occur at Graffham Common. Perhaps buxi was closest but that was a long way away in London and given that there were two records of emarginatus from the exact same 10 km square, that seemed most likely. Occam's Razor and all that.

I fed the little fella up on springtails from leaf litter in the garden and it did really well, surprisingly reaching adulthood today. The frustrating thing is that the left palp did not come out of the skin when it last shed its skin (it shed its skin twice by the way). So here are some more shots of the nationally rare and Vulnerable Philodromus emarginatus, the 392nd spider I saw last year but the one I can't count either on last year's list, or this year's. Man, this is one gorgeous spider.


Persistence always pays off. Looking at the SRS page it's not been recorded since 2015 and Mark Gurney says the only other records for a National Trust site were from Lavington in West Sussex where it was last seen in 1996. The last Hampshire and New Forest record was from the same 10 km square in 1999. Very cool and the only good thing about the 31st January 2020. Now I wish I had got that female Enoplognatha from Cornwall to adulthood all those years ago but that's another story...

Rare Spider Not Seen this Century Found in Black Hole

Posted by Graeme Lyons , Sunday, 19 January 2020 08:36

Before I found myself sucked into the inescapable gravitational pull of Black Hole at Burton Pond, I headed to Iping Common to MAX OUT ON MY SPIDER YEAR LIST! There, I've said it. I'm year-listing spiders again apparently. Anyway yesterday was awesome. From Iping and Burton I recorded 65 species of spider which from mid winter on a frosty day I think is pretty smart. I got TWO lifers out of it. Neither were from Iping. Iping was great for some big impressive spiders I would normally expect to see in autumn, such as an adult Ero tuberculata above (looks like another new record for January). This adult female Agroeca proxima

Other charismatic mega fauna included Marpissa muscosa.

Nuctenea umbratica. I don't think I have ever noticed their weird eye arrangement before.

Clubiona corticalis with the above two on the same tree.

And a Cream-streaked Ladybird just to keep my eye in with the beetles.

I had collected tons of linys which it turned out were going to be mostly unexciting species. The Sphagnum areas at Iping were yielding nothing so by about 1.30 pm I found myself sucked into the Black Hole at Burton Pond.

A year on I was very much aware how much my spider field craft, especially with money spiders, had moved on. I was ticking off things such a Sintula corniger, Taranuncus setosus, even female Notioscopus sarcinatus (I did check these). In addition immature Theridiosoma gemmosum and Rugathodes instabilis. Black Hole really is a gem and recent work that Jane Willmott has done there is clearly having a great benefit to the site by keeping it more open. An an adult Platnickina tincta was a bit of a surprise too.

Oh and Demetrias monostigma. If you are finding this beetle, you are on a good site for spiders. And beetles. I only ever see it in old Greater Tussock-sedge and Marram tussocks.

It wasn't until I got home though that I realised I had found something rather good.

Here is Centromerus brevivulvatus. Last seen in the UK in West Sussex in 1998 by Dick Jones at Durford Heath. It's Endangered and Nationally Rare. Only ever known from seven 10 km squares, four of which are in Sussex. Interestingly one of these is Iping Common where it was last seen in 1969. This record though is an entirely new 10 km square making eight 10 km squares, five of which are in Sussex! Oddly only this week I assessed this species as a 'Sussex rare species' and thought to myself "I should get out there and have a look at Durford Heath."

Have a look at the SRS page for it here. Here is the map. Look at all those 'x's.

The singularity that is the Black Hole opened up an Einstein-Rosen Bridge to 1998. What was I doing then? This:

I didn't know anything about spiders then, but I did look a lot more like a spider than I do now. I'd have been half way through my astrophysics degree at Sussex Uni at this point (that's not the name of a hipster ale by the way {hipsters didn't exist then} - in the above image I am halfway through a pint of Hoegaarden).

Anyway, back to Black Hole. One of the next spiders I looked at I only recently ticked off up north. I was amazed to find a male Aphileta misera. This seems to be a species new to West Sussex, the last Sussex record from Ashdown Forest was in 1970. So where was I in 1970? Well I was an unconstituted smear of matter across the Staffordshire landscape, with smaller clusters peppered around the world depending on where my mother's food would come from some seven years later. Some of my carbon might even have been sequestered by spiders at that point, probably the bits that would one day make up my neurons. We have all been there!

And also new for me was Diplocephalus permixtus. A species I was expecting to see up north. Turns out this is the first West Sussex record since 1944 and Sussex record since 1999.

Now I did quite a bit of recording at both these sites last year and it really does go to show how important repeat visits are. With just shy of 10% of the UK fauna seen in a single day I will not be doing any recording now until next weekend.

Hot stuff

Posted by Graeme Lyons , Thursday, 16 January 2020 21:50

This story ends with a spider in a butterfly house in a zoo and starts with a phone call a week earlier about pan-species listing. There is a thrush in the middle.

On Monday I went up to see Mark Telfer about buying a microscope off him after discovering quite by chance he was selling one. No point driving 100 miles to see an old friend without some natural history though, so we were soon heading over to Whipsnade Zoo. A month after the Black-throated Thrush was first seen there. I like to think of this as 'danger twitching', waiting as long as you possibly can before going for the bird. It was an effortless twitch and I took this absolutely stunning photo. My first new British bird since the Elegant Tern in 2016. This is a really awesome bird and deserves more than this but this just isn't his story.

Then we went into the butterfly house. Now I have done very little in hothouses, my only attempt was at the BMIG excursion to Wisley Garden Centre. I was a little bewildered by how many species Mark was finding, such as these impressive molluscs which Mark identified as Subulina octona. My completely steamed up camera and hand lens were making me feel out of my depth.

The spiders seemed a bit more do-able. A single large Ulobrous plumipes and dozens of Achaearanea tepidariorum (with some adults of both sexes) were present along with a few familiar Pholcus phalangiodes. Mark picked up a couple of little orange things that looked very like Oonops which I suggested they were. I couldn't find one beyond a very small immature. Then I noticed something scrambling around in the leaf litter. Possibly a parasitic wasp, I just about got it into a tube to see it was a tiny ant mimic type thing that was clearly an adult male. I vaguely remembered there being something of this shape in the new book and a quick look through and I thought I wouldn't be far off calling this Coleosoma floridanum.

I was very excited to get home and see it hadn't been seen since 2005. What a strange beast, hard to believe it's a theridiid. This animal was just over 2 mm long. It has a strange peg on the underside of the abdomen, two serrated projections at the front of the abdomen, a strange cleft in the top and two undulations underneath. 


Yet the fun didn't stop there, turns out the Oonops type things were not one of our two Oonops. Mark thinks it could possibly be Triaeris stenaspis. The really odd looking abdomen and long patella of the first leg would add up. It's turned up in hothouses in Europe and apparently at some point in the UK. Thanks to Mark for this research, image and an awesome day! I do think it would be great if the SRS had distribution maps for all spiders that are surviving in places like this in the UK. They might not be able to survive outside but they are very much breeding here and not under anyone's control (i.e. they are not pets). Are they really any different to Uloborus plumipes? I can see the appeal now and I think I will be doing some more hothouses soon!

EDITS: A second update from Mark: "T. stenaspis was recorded from the Eden Project by Snazell, R. and Smithers, P. and published in Bull BAS in 2007, referring to sampling carried out between 2002 and 2004". My previous edit that this was only the 2nd UK record since 1909 is now known to be incorrect. 

Which leaves one question: Hypothetically speaking, if one was year-listing spiders, would this species count on one's list? Hypothetically speaking obviously.
(One is on 38 species).

My 1000th post!

Posted by Graeme Lyons , Sunday, 12 January 2020 19:19


*I AM DEFINITELY NOT YEAR LISTING SPIDERS*.

I noticed just the other day that my next post would be my 1000th! I have nearly been blogging for ten years and blocking up people's feeds with gibberish about wildlife for so long I can't remember not doing it. DID I used to NOT write this blog? I can't be sure. Can anyone remember not reading it? 

I had an interesting day doing some casual recording at Castle Hill on the edge of Brighton in my home 10 km square today. It's the first time I have gone out with my suction sampler for ages, well for 12 days to be precise. An electric suction-sampler is such a great way to terminate boring conversation in the field, just turn it on and instantly your friend's incessant monologues are drowned out as you collect a hat full of spiders! It's the gift that keeps on giving. (Carole, I only wrote this because you said my blogs used to be funny, I didn't mean it. I could still hear you).

I was trying not to focus just on spiders but I kind of found myself focusing rather a lot just on spiders. Carole Mortimer was keen to spend some time at an NNR with me recording spiders so we did just that and I managed 28 species, 18 of which were new to the 10 km square. I feel quite embarrassed by that as I live in it and did quite a bit of spider recording last year. The species above is the nationally scarce/S.41 Ozyptila nigrita. It's fairly regular on the chalk in the summer but I have never seen it in January before as an adult, no one has it would seem looking at the scheme. Both male and female were present today. I didn't see this until mid April last year. These warm winters are odd.

I took a  load of linys home and amazingly got a lifer! So a species I clearly didn't see last year. A female Diplocephalus picinus. Not scarce but clearly not common in Sussex.

And earlier, another spider that I have only recorded twice, both at 450 m in the Peak District and never in Sussex. The first Sussex record of Poeciloneta variegata since 2007 (and the first East Sussex one since 2004).

Here is today's list with those in bold new to the 10 km square and cons statuses in brackets. The four scarce things are all good chalk-grassland species.

Agalenatea redii
Centromerita concinna
Cnephalocotes obscurus
Diplocephalus picinus
Ero cambridgei
Euophrys frontalis
Gonatium rubens
Hypsosinga albovittata (NS)
Hypsosinga pygmaea
Mangora acalypha
Meioneta rurestris
Meioneta simplicitarsis (NS)
Micrargus herbigradus
Microlinyphia pusilla
Neottiura bimaculata
Ozyptila brevipes
Ozyptila nigrita (NS, S.41)
Pachygnatha degeeri
Palliduphantes ericaeus
Panamomops sulcifrons (NS)
Pelecopsis parallela
Pholcomma gibbum
Pisaura mirabilis
Poeciloneta variegata
Selimus vittatus
Tenuiphantes tenuis
Zora spinimana
Zygiella atrica

So if you make a site list of spiders (28), add it to the spiders you have recorded in your flat this year (3), apparently that's year-listing spiders but as I said earlier, I AM DEFINITELY NOT YEAR-LISTING SPIDERS.

I end the day on 31 spiders for the year 

(BUT I AM NOT YEAR-LISTING SPIDERS, THAT'S JUST THE TOTAL OF ALL THE SPECIES OF SPIDER I HAVE SEEN THIS YEAR).

My top ten spider highlights of 2019 and the full species list

Posted by Graeme Lyons , Saturday, 4 January 2020 18:18

I recorded 391 species of spider in 2019 and 88 of these were lifers. December proved to be the best month of all where I added 42 species and 38 of these were lifers! 21 lifers in Staffs and 16 on the trip to the West Country. Here are my top ten spider highlights in reverse order:

11. My first ever Singa hamata
I had literally just met a new client at a site in Hampshire and in the first sweep net full was this Singa hamata. Being much larger than I expected it to be and not remotely on my radar for the site, I squealed "What the @*&! is that!". Turns out it's the first Hampshire record since 1991.

10. Rye Harbour
It was great meeting up with Matt, Chris and others at Rye Harbour, Camber Sands and Castle Water and we added shed loads of species to our year lists in the time we were there. Such as this female Pellenes tripunctatus.

9. More and more rare spiders recorded at Graffham Common
I pushed up the total number of species recorded on Sussex Wildlife Trust's reserves to exactly 400 species by the end of 2019 and Graffham Common shines out as a brilliant spider site, including probably the best place I know to find Araniella displicata.

8. Cementing Iping Common as the best site in Sussex for spiders 
With 220 species and 55 with conservation status, Iping Common is confidently the top spider site in Sussex. Probably the best day there was in spring when I recorded Mecopisthes peusi (last recorded there in 1968 and in Sussex in 1989). Additionally, I also recorded Tapinocyba mitis on the same day in late February. It was around this point that I had started throwing some time and energy into year listing spiders and was really getting into it.

7. East Head invertebrate survey
The spiders on this sand dune site in West Sussex are just amazing but this little chap stole my heart. Sitticus saltator you are the best.

6. Stumbling across Haplodrassus silvestris on a woodland ride
Probably the most unexpected spider of the year. I was heading rapidly across a site to do a survey and saw what I thought was a Scotophaeus blackwalli. It was actually the first Sussex Haplodrassus silvestris since 1908.

5. Finding Scotina palliardii twice in West Sussex. 
A single animal at Levin Down on my birthday and 16+ at Kingley Vale in mid Deecmber. The first Sussex records for some time with only one known record with a vague date for Shoreham.

4. A day out in Devon
Meeting up with Matt for only the second time in 2019 but this time on his turf got me a shed load of ticks and it was great to meet up with John Walters too who took us right up to four Nothophantes horridus.

3. Kynance Cove
Tylan Berry took me to Kynance Cove on the Lizard and we recorded five national rarities in an hour and four of these were new to me, I think the highlight for me was finding an adult female Segestria bavarica under the second rock I turned over but the rarest would be the Gnaphosa occidentalis.

2. Philodromus fallax!!!
After years of searching for it in a range of different sand dunes, it popped out of a Marram tussock in early November just as I was leaving Camber Sands in East Sussex! What a beauty.

1.The Staffordshire Hoard
I couldn't have done it without the big twist at the end of the year. I went on holiday by mistake. Didn't think my final move would be quite as successful as it was but the trip to the Roaches where I had five lifers (and the following day at Gun Moor where I had another six) was awesome and without these two days specifically it would have been a different result. Finding some real upland species like this Porrhomma montanum under the first rock I looked at was really lucky up the Roaches. I can't believe I just said my highlight of the year was a Porrhomma. It was the whole experience, not just this spider though. The Roaches are just breathtaking. Schwingmoor is so much fun and I can't get enough of all that Sphagnum. Low cloud can do one though.

So there we have it. The winter money spiders were the thing that did it for me the most. Anyway, as promised, here are all 391 species in the order I recorded them (approximately - those at the start are a bit random, maybe the first 60 or so species). A total of 88 lifers or 22.5% were new species for me! Of the 391 species, 315 were in Sussex and 261 were on Sussex Wildlife Trust reserves.

I have totally levelled up with spiders this year and this is the key point to remember, the winners here are the spiders and spider recording. It's great to see so many people listing spiders this year now but I won't be, I need a break. Nice to have a record to smash, I am sure someone will get 400 soon. Massive thanks to Matt for starting the whole thing. Here is my full list. Lifers in bold. Typos expected. Look how the lifers stack up at the end!

Order Species Status
1 Pisaura mirabilis
2 Pholcus phalangioides
3 Steatoda nobilis
4 Erigone atra
5 Meioneta mollis NR
6 Neon reticulatus
7 Diaea dorsata
8 Gongylidium rufipes
9 Anelosimus vittatus
10 Coelotes atropos
11 Steatoda grossa
12 Pholcomma gibbum
13 Phrurolithus festivus
14 Harpactea hombergi
15 Tenuiphantes tenebricola
16 Ero aphana NS
17 Macoreris nidicolens
18 Megalepthythantes nr. collinus
19 Bathyphantes gracilis
20 Tenuiphantes zimmermani
21 Palliduphantes ericaeus
22 Tenuiphantes mengei
23 Crustulina guttata
24 Cercidia prominens NS
25 Zora spinimana
26 Philodromus aureolus
27 Pseudeuophrys lanigera
28 Segestria senoculata
29 Alopecosa barbipes
30 Pachygnatha clerki
31 Psilochorus simoni
32 Xygiella x-notata
33 Kochiura aulicus NS
34 Mangora acalypha
35 Simitidion simile
36 Zilla diodia
37 Agalenetea redii
38 Hypsosinga pygmaea
39 Pachygnatha degeeri
40 Pardosa nigriceps
41 Episinus angulatus
42 Lathys humilis
43 Talavera petrensis NR
44 Hypsosinga sanguina
45 Hypsosinga albomaculata NS
46 Euophrys frontalis
47 Episinus truncatus NS
48 Philodromus histrio NS
49 Pardosa saltans
50 Lophomma punctatum
51 Clubiona lutescens
52 Cnephalocotes obscurus
53 Savignya frontata
54 Pepenocranium ludicrum
55 Meioneta rurestris
56 Kaestneria pullata
57 Micrargus herbigradus
58 Gonatium rubens
59 Mecopisthes peusi NS
60 Tapinocyba mitis NR
61 Micrommata virescens NS
62 Hypselestes jacksoni NS
63 Ozyptila atomaria
64 Ero cambridgei
65 Notioscopus sarcinatus NS
66 Oedothorax gibbosus
67 Taranuncus setosus NS
68 Clubiona subtilis
69 Nuctenea umbratica
70 Marpissa muscosa NS
71 Metellina segmentata
72 Hahnia nava
73 Minyorilus pusillus
74 Gibbaranea gibbosa
75 Metopobactrus prominulus
76 Amaurobius fenestralis
77 Cyclosa conica
78 Larinioides cornutus
79 Tenuiphantes tenuis
80 Neottiura bimaculaa
81 Microneta varia
82 Oedothorax apicatus
83 Tegenaria gigantea
84 Hypomma bitubermaculatum
85 Amaurobius similis
86 Coelotes terrestris NS
87 Uloborus plumipes
88 Labulla thoracica
89 Lepthyphantes minutus
90 Clubiona corticalis
91 Araniella displicata NR
92 Ozyptila trux
93 Tenuiphantes flavipes
94 Ozyptila brevipes
95 Gnanthonarium dentatum
96 Ero furcata
97 Walckenaria nudipalpis
98 Porrhomma pygmaeum
99 Walckenaeria unicornis
100 Silomeptus ambiguus NS
101 Halorates reprobus NS
102 Tiso vagans
103 Xysticus cristatus
104 Neriene clathrata
105 Sintula corniger NS
106 Salticus scenicus
107 Theridiosoma gemmosum NS
108 Arctosa perita
109 Microlinyphia pusilla
110 Diplostya concolor
111 Oedothorax retusus
112 Bathyphantes parvulus
113 Clubiona phragmitis
114 Clubiona stagnatalis
115 Ceratinopsis stativa NS
116 Amaurobius ferox
117 Dysdera croccata
118 Araneus diadematus
119 Alopecosa cuneata NS
120 Bathyphantes approximatus
121 Enoplagnatha thoracica
122 Erigone dentipalpis
123 Euryopis flavomaculata NS
124 Araneus triguttatus
125 Walckernaria antica
126 Pardosa pullata
127 Pardosa paludicola NR
128 Dipoena inornata NS
129 Pelecopsis parallella
130 Sibianor aurocinctus NS
131 Dictyna arundinacea
132 Padiscura pallens
133 Clubiona compta
134 Neriene peltata
135 Textrix denticulata
136 Trochosa ruricola
137 Clubiona diversa
138 Panamomops sulcifrons NS
139 Talavera aequipes
140 Drasyllus pusillus
141 Xysticus kochi
142 Phlegra fasciculata NR
143 Sitticus saltator NS
144 Stemonyphantes lineatus
145 Crustulina sticta NS
146 Ceratinopsis romana NR
147 Pardosa proxima NS
148 Zelotes electus NS
149 Walckenaeria vigilax
150 Cheiracanthium erraticum
151 Anyphaenea accentuata
152 Trematocephalus cristatus NS
153 Pardosa prativaga
154 Alopecosa pulverulenta
155 Ozyptila sanctuaria
156 Micaria pulicaria
157 Trochosa terricola
158 Xysticus audax
159 Xerolycosa miniata NS
160 Thanatus striatus NS
161 Agelena labyrinthica
162 Trachyzelotes pedestris
163 Monocephalus fuscipes
164 Clubiona terrestris
165 Xysticus bifasciatus NS
166 Phaecodeus braccatus NR
167 Theridion mystaceum
168 Pardosa monticola
169 Hybocoptus decolatus NS
170 Xerolycosa nemoralis NS
171 Evarcha arcuata NS
172 Hypomma cornutum
173 Antistea elegens
174 Zelotes latreilli
175 Neriene montana
176 Episinus maculipes NS
177 Dicymbium nigrum
178 Misumena vatia
179 Ballus chalybeius NS
180 Mermmessus trilobatus
181 Pardosa agrestis NS
182 Myrmarachne formicaria NS
183 Platnickina tincta
184 Heliophanus flavipes
185 Diplocephalus cristsatus
186 Argenna subnigra NS
187 Pirata piraticus
188 Dolomedes plantarius NR
189 Argyroneta aquatica
190 Arctosa leopardus
191 Podacinemis juncea
192 Pardosa amentata
193 Scotina palliardii NR
194 Zodarion italicum NS
195 Xysticus lanio
196 Cicurina cicur NS
197 Hylyphantes graminicola
198 Linyphia hortensis
199 Theridion impressum
200 Robertus arundineti (26th April)
201 Dictyna uncinata
202 Ozyptila nigrita NS
203 Xysticus erraticus
204 Neoscona adianta
205 Neon pictus NR
206 Theridion melanurum
207 Haplodrassus signifer
208 Philodromus albidus
209 Pardosa palustris
210 Erigonella hiemalis
211 Araneus quadratus
212 Singa hamata NS
213 Maso sundavelli
214 Pachygnatha listeri
215 Araniella curcurbitina
216 Araniella opisthographa
217 Ozyptila simplex
218 Metellina merianae
219 Osterius melanopygius NS
220 Argenna patula NS
221 Enoplognatha mordax NS
222 Heliophanus cupreus
223 Clubiona reclusa
224 Theridion sisyphium
225 Erigone arctica
226 Pardosa purbeckensis
227 Drassyllus lutetianus NS
228 Pirata latitans
229 Dolomedes fimbriatus NS
230 Philodromus rufus
231 Xysticus ulmi
232 Theridion pictum
233 Xysticus acerbus NR
234 Tibellus oblongus
235 Pirata hygrophilus
236 Philodromus dispar
237 Theridion blackwalli NS
238 Uloborus walckenaeria NR
239 Evarcha falcata
240 Philodromus margaritatus NR
241 Pocadicnemis pusillus
242 Araneus sturmi
243 Pirata uliginosus
244 Kaestneria dorsalis
245 Dipoena prona NR
246 Maso gallica NS
247 Tetragnatha montana
248 Erigone promiscua
249 Tetragnatha extensa
250 Clubiona brevipes
251 Clubiona pallidula
252 Aellurillus v-insignitis NS
253 Gongylidiellum vivum
254 Neriene furtiva NS
255 Haplodrassus silvestris NS
256 Theridion varians
257 Pellenes tripunctatus NR
258 Trichopterna cito NR
259 Drassodes lapidosus
260 Trichoncus affinis NR
261 Marpissa nivoyi NS
262 Argiope brunnechi
263 Cheiracanthium virescens NS
264 Haplodrassus dalmatiensis NS
265 Lathys stigmatissima NR
266 Drassodes cupreus
267 Tetragnatha obtusa
268 Anyphaena sabina
269 Nigma puella NS
270 Philodromus collinus
271 Dictyna latens
272 Rugathodes instabilis NS
273 Philodromus praedatus
274 Araneus angulatus NS
275 Steatoda phalarata
276 Zygiella atrcia
277 Achaeranea simulans
278 Enoplognatha ovata
279 Sitticus pubescens
280 Philodromus cespitum
281 Ozyptila praticola
282 Linyphia triangularis
283 Drapetisca socialis
284 Theridion pinastri NS
285 Dipoena erythropus NR
286 Scotina celans NS
287 Steatoda bipunctata
288 Agyneta cauta NS
289 Dipoena tristis NS
290 Enoplognatha latimana
291 Hahnia montana
292 Erigone longipalpis
293 Tetragnatha nigrita
294 Ero tuberculata NS
295 Sitticus inexpectus NS
296 Thomisus onustus NS
297 Scotina gracilipes NS
298 Araneus alsine NS
299 Achaeranea lunata
300 Agroeca inopina
301 Agroeca proxima
302 Oonops domesticus
303 Salticus cingulatus
304 Scytodes thoracica
305 Nigma walckenaeria
306 Walckneaeria acuminata
307 Helophorus insignis
308 Gonatium rubellum
309 Ceratinella brevipes
310 Robertus neglectus NS
311 Larinioides scloptarius
312 Tetragnatha striata
313 Segestria florentina
314 Araeoncus humilis
315 Centromerita concinna
316 Araneus marmoreus
317 Floronia bucculenta
318 Diplocephalus latifrons
319 Tenuiphantes cristatus
320 Centromerus dilutus
321 Walckenaeria cuspidata
322 Philodromus buxi
323 Typhochrestus digitatus NS
324 Pirata piscatorius NS
325 Pelecopsis nemoraloides NS
326 Dysdera erythrina
327 Clubiona trivialis
328 Walckenaeria monocerus NS
329 Agroeca cuprea NR
330 Philodromus fallax NR
331 Ceratinella brevis
332 Palliduphantes pallidus
333 Agroeca brunnea
334 Tapinopa longidens
335 Hahnia helveola
336 Meta menardi
337 Centromerus arcanus
338 Meioneta simplicitarsis NS
339 Lepthyphantes leprosus
340 Porhomma microthphalmum
341 Nesticus cellulanus
342 Macrargus rufus
343 Robertus lividus
344 Oedothorax fuscus
345 Tallusia experta
346 Metellina mengei
347 Monocephalus castaneipes NS
348 Centromerus sylvaticus
349 Centromerita bicolor
350 Walcknaeria nodosa NS
351 Tenuiphantes alacris
352 Nothophantes horridus NR
353 Cryptachea blattea
354 Tegeneria saeva
355 Thyreosthenius parasiticus
356 Pelecopsis sussanae
357 Clubiona genevensis NR
358 Gnaphosa occidentalis NR
359 Euophrys herbigrada NR
360 Segestria bavarica NR
361 Achaearanea tepidariorum
362 Cryphoeca silvicola
363 Tegenaria silvestris
364 Gongylidiellum latebricola NS
365 Centromerus serratus NR
366 Walckenaria alticeps NS
367 Porrhomma pallidum
368 Drassyllus praeficus NS
369 Scotophaeus blackwalli
370 Dicymbium tibiale
371 Erigonella ignobilis NS
372 Bolyphantes luteolus
373 Poeciloneta globosa
374 Centromerus prudens
375 Porrhomma montanum NS
376 Scotionyrilus evansi NS
377 Saaristoa abnormis
378 Theonoe minutissima
379 Micrargus apertus
380 Tapinocyba pallens
381 Aphileta misera
382 Bolyphantes alticeps
383 Oonops pulcher
384 Saaristoa firma NS
385 Bathyphantes nigrinus
386 Latithorax faustus NS
387 Agyneta decora
388 Oryphantes angulatus NS
389 Hilaira excisa
390 Pelecopsis mengei
391 Gongylidiellum murcidum NS

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